Lag0003227.1 (mRNA) Sponge gourd (AG‐4) v1

Overview
NameLag0003227.1
TypemRNA
OrganismLuffa acutangula (Sponge gourd (AG‐4) v1)
DescriptionH(+)-exporting diphosphatase
Locationchr4: 49239942 .. 49246867 (-)
Sequence length327
RNA-Seq ExpressionLag0003227.1
SyntenyLag0003227.1
Sequences
The following sequences are available for this feature:

Gene sequence (with intron)

Legend: polypeptideexonCDS
Hold the cursor over a type above to highlight its positions in the sequence below.
ATGAGACAGATTCTCAAGGCCTCCGCTGTGGGCAGCATAATGTATGCAATGCTCTGCACAAGGCCTGACATCTGTGATGATGTAGGGATAGTTAGCACATATCAATCAGTATCCATGGTCAAACTTTACAAAGGAGCTATAATGTGGCATAATTACAGTCTTAACCATATAAGCCAAAAATATAGTTGCTTGTGAAGCAGCGATAGAGATTGTAAGGTTTAGAAAGTTCTTGACTGATTTGGAATGTGTTCCAAATGTGAACTTGCCCATCAGACTTTATAGTGATACAGTGGTGTTGTGGCAAATTCATCAGAACTTTCCAGCCCAAGACAAGCTCACACATTGAATTAAATTGAAATATTCACGAAGACTCTTTCGGCTAAAGGGTATGAGGGTCATCTAGAAGGTCTAGGTCTACGAGTTATGTACAAAGATGAAATCTGGGGCAAGTGGGAGAAATGTAATGGGTATAGTGATATCATCATACATTGGGTATAGTGATGCCCTAGTTTATTGTATTTGTACATTGTATATGTTTTATACATGTATTTATACACCCACTATTGAATAAGATTGTTGTACATCCCACTAAACTAGAGTTTTAGTTAAAGTGGGAGTTTGTTGGGTTTTATGCCCTAAAACTCGTAGATAGTGAATGTAAACAAATTGTATACTTGTATTTAATAAAGTGTTATTTAATTTTCTGCAATAAAATGTTATTGATAAATATTGCACTATTTGACCCTAATCCAATAAACTAGACATCCTAGGCTGTAATATGAGTCTTGAACTATATGTAGCGACATATGGGGATCATTGTTCAAGAAACAACCTAAAAGGTCTATAGTATAGGGATAAGACTGGGTGCCTTATCCTGGCAACACTATGGATACGACCCACTTTGTATTTGATACAAACGCTGCGATCCAACGCGTTCGTGTAGGAGACATGCGAGTTGAGGTATCCTATGCAATGAGTTTGCATAAGACCAGACCGCGAAATAGTAACCACTGGTTATAACACCGTTAACTAGTTGGTTTTCTATTTCACTAGGACGACCTAGACAACTTAGTCTTAATCCTGAGTTGATTATGGACTCCTGCTCATGAGAGATTGTCCTTTGATTTGTATGGATGAGAGTGGCCAATACGCCGACTCAATAAGCCTACCACTTTGGGGACAAGACCGAATGAGGAGCTGGGAACATAATCGTACAAGATGGAATTCACACCTTCCCGACTTTAGGGAAGCAGATGAGTGTTCCCTTAAGTGGTTACTCCGAGTCTTGAACAGAGGGCCTTACCCTCTCAATGGCACGAGAGGGTTTTCTGTTTGATGGTTGGACCACAAACAAGTTGTTCATTAGAGGAGAATTGGTACTTAAGGAAATAGAGGTAACCCAGGTTTCCTCTTTTCTTTAGAAAAGTTTTAAAAATAAATTTAGATTTATTTTAGCATGTTTAAACCAAATTTTACATCGGCTTTAAAGGTTGTTTAAAGTTTTGTTTGAGTTATCGTAACGGTATTTTAATGTTAATTAATATACGGCGTAAGGTCCAATCGATGCGCTTCCGCTGCGTGAGGACTTTTATTCCCTTCACATAATTGTACCTATAAAAGGATGAACTTCCATGTAAAGTTTGCAACACCAAATTTTCTACTTCAATTAGAAGTTTATTCATCTTTTGGATTTCACACTCTCTCGTTTTCTATTTTCTCTCTTTGTTCAACAAACATTGAGAGTTTGATTATTGCTTCCGCAATATTTTTTCGACCTTTCAATTTTCAATTTTCTCAACATTACCTTATAAAAAATAAAAATAATAATAAACAATACGAAATTATACTAACAATTAATTTTAAAAAGTCAAGTAGTTTTTAAAATCAAGTAAGTTACATGTTAAATTGCCTTTTCTAAGATTAAAAAAAAATATAATATAAGATTTAAGCAAAACAATTTATTATTAAAATTTAGGGCTGGTTGCAAGGATAGCAATATGGGCCAAAATAATTGCAAATCTAGCAACCCACAAAAAAAATTGCAAAAATGGCAAAATCTAATAGTTTGTGTGTTGCATTTACTAAAATACCCCTCCAGTTTTTCTTTCACTAATAGAAATTTTTTCTATCACTGATAGATTCTATCAGTGATAGAAAGATTTTCTATCAGTGATAGAAATTTTCTATCACTGATAGAAAGATTTTCTATCATGATAGAAATTTTCTATCAATGATAGATTCTATCAGTGATAGAAATTTTCTTTCACGGATAGATTCTATCATATGATAGAAAATTTTTCTACTACTGATAGATTCTATCGGCGATAGATATTTTTCTATCACTTGTTCGATGTTCTATCAGTGTTTGGTTGTTGTTATTTGGTTTTCTATCAGTGTTTGGTTTTTGCTTTTCAGTGTTTGATTTCTATAATCGTTCGTTTTTGCTTTATTTTTCAATGCTTGATTTCACCATGTATCACCATCGCTGATAGAAACAAAAGATCTTCTATCACCGATAGTAAAGTATCAGTGATAGAAAGATTTTCTATAACTCACGAAAAGTTTTTCTTTCAGTCTCAGTGATAGAGATAGAAAGTTTTTCTATCACTCATAGAAATATTTTCTATGAATGATAGAGATTTTTTCTATCACTCATAGAAAAAATTTCTATAAGTGATAGAAGATTTCTATCACTAATAGGTTCTATCAGGGATTGAAAGATTTTCTATCAAGAGTTTTCTATCATTGATAGAATATCTTTCTATCACTGATAGAATAGTAAAAATTCTATCAGTAAAAGATCGATAGAAAGCTTTTCTATCGGTGATAGAAAGCTTTTCTATCGGTGATAGAAAGTTTTTCTATCACTTATAGATTATATCGGTGATAGAAAGATTTCTATCGGAGATAGAAAGATTTTTATGGATGATCAGGGATTGAAAGATTTTTTATCAAAAGTTTTTCTATGATTTTCTATAAAAAGTTTTTTTATCACTGATATAACCTATCACTGATAGAAATCTTTTATCACTGATAGAAAAAGTTTCTATTAGTGATAGAAATCTTCTATTCTATCAGTGATAGAAACTTTCTATCACTGATAGAAAATTTTTCTACCACTGATAGAATCTATCAGTCATAGAAATATTAGTGATAGAAACTTTCTATCGCCGATAGAAAATTTTTCTACCACTGATAGAATCTATCAGTGATAGAAATATCACTGATAGAAACTGTATTCTATCAGTGATATAAACTTTCTATCACTAATAGAAACTTTATTCTATCAGTGATAGAAACTTTCTATCACTGATAGAAAATTTTTCTACCACTGATAGAATCTATCAGTAATAAAAAGACTTTCTATCAGTAAAAAAAAAAAAAAAAAAAAAAAAAAAAAAACTGACGAGGAAGAAGAAGATGGGCACACAGGGGCAATTTAGGAATATAAAAAATTAATTAAAAAATTTTGCCATATTTGCAAATATTTTTAAAAATTGCTATATTTGTAATTATTTACCTTCATGTTGCCACCCATTACAATTATCCTAAAATTTAAGTAGCTGATGTTAGATAATTAATAGCATTAAAAAACTTTCCACTAAAACCAATTACATAAATTTAAATATAAATACTAAAAGTTCTTGTCTTTGTGATTAATGAACAAAAAATTAATCTATTCAATTAGAAAATTGAGAGTGTTTTCTCTTCAAAAAAAAAAAAAAAATAATAATAATAATAATAATAATTGAGAGTGTGCTCAATTATTTTGGTGTTTGAAAATTTTAATTTCATTTAAAATTATGCATGCATTTATTAATTAATAAGTTTTTTTTAAGGAATAACTTAATAATTTTTAGGCTACTGAGTTTTATTTTTATTTTATTATTATCATTTTCACACTTATAAGTTTCCAACCTAAAATGTATTAATTAGTTTGTATATTAAAATACAAATTTGGTCATACATTTAGACAATTTTATTTTGGTCACTGTGGTTTTAAAAGTTTCAATTTAGTCCCAATAGTTTGAGGATAACTTCAATTTTCTTCCTATGGTTTGGATTTAGTTCCAATTTGTCTAAGTCATCCCTAAATTGTATTTTACCAACTTGAAAGTCCACGTTTTTTCTTTTTATTTTTACCTAGCCCTAAACTCCAAGTTAGTCAACTTAACACAAATTACTTAGATAGAAACTTGCTACATAAAAAAAATGGCAATATATGATAACGACAAAGAAAACTACAATAATGTAATAATCAAAATAGAAACATGCTCATAAATAAAATAAGTGATTTTCTTACGGATGACCCTCTAAAATAGACGAAAATGATAAATGTCCTACATTTTCAATTGGATATATGTCATTTTGCTTGCGTAAGCATCTTAGTAAAATTACCATTCTACTCTCTACTCTAACCTCGTTCACTTTCGCTACTCTTGCTCTCTCTCCAACCCTTTTTTTTTTTTAAAAACAGTGGAAGAAGAAAAGGGAGGGATGAAGAGGGAAGACAAATAAGGAAAAAAGGAGGAGAAGGAAGAAGAATAAAAGAGGAGAAAAATAGAAAAAAAATGAAAAACGACAGTTTTCGGCGGCGGTATCGACAACGACGGTGGCTACTGCAAATGAATTGAGATTAGGGTTTGAAGGAGAGGAGAAGAAGATTAAGGAAGAAAATGGAAGAAGAAAAAAGAAAAGAAAAAAAATAATAAAAAAGGATTTAATATAAGGTCGGTTGCAAGGATAGCAATATGGGCCAAAATAATTGCAAATCTAGCAACCCACAAAAAAAAATTGCAAAAATGGCAAAATCTAATAGTTTGTGTGTTGCATTTACTAAAATACCCCTCCAGATTTTCTTTCACTGATAGAAAGTTTTTCTATCACTGATAGATTCTATCAGTGATAGAAAGATTTTCTATCAGTGATAGAAATTTTCTATCACTGATAGATTCTATCAGTGATAGAAATTTTCTTTCACGGATAGATTCTATCATATGATAGAAAATTTTTCTACTACTGATAGATTCAATCGGCGATAGATATTTTTCTATCACTTGTTCGGTGTCTATCAATGTTTGGTTGTTGTTATTTGGTTTTCTATCAGTGTTTGGTTGTTGTTATTTGGTTTTCTATCAGTGTTTGGTTTTTGTTTTTCGGTGTTTGATTTCTATAATCGTTCGTTTTTGCTTTATTTTTCAATGCTCGATTTCACCATGTATCACCATCGCTGATAGAAACAAAAGATCTTCTATCACTGATAGAAAAGTATCAGTGATAGAAAGATTTTCTATAACTCACGAAAAGTTAGGTTCTATCAGGGATTGAAATATTTTCTATCAAGAGTTTTTTATCATTGATAGAATATCTTTCTATCACTGATAGAATAGTAAAAATTTCTATCAGTAATAGAAATCTTTTATCACTGATAGAAAGCTTTTCTATCGGTGATAGAAAGTTTTTCTATCACTTATAGATTATATCGGTGATAGAAATATTTTTATGGATGATCAGGGATTGAAAGATTTTTTATCAAAAGTTTTTCTATGATTTTCTATAAAAAGTTTTTTTTTATCACTGATAGAACCTATCAGTGATAAAAATCTTTTATCACTGATAGAAAAAGTTTCTATTAGTGATAGAAATCTTCTATTCTATCAGTGATAGAAACTTTCTATCACTGATAGAAAATTTTTCTACCACTGATAGAATCTATCAGTCATAGAAATATTAGTGATAGAAACTTTCTATCGTCAATAGAAAATTTTTCTACCACTGATAGAATCTATCAGTGATAGAAATATCACTGATAGAAATTTTATTCTATCAGTGATAGAAACTTTCTATCACTGATAGAAAAATTTTCTACCACTGATAGAATCTATCAGTAATAAAAAGACTTTCTATCAGTAAATAATAATAAAAAAAAATTAAAAAAAAACAAAAAAAAAAACTGACGAGAAAGAAGAAGATGGGCACACATGGGCAATTTAGGAATATCAAAAATTAATTAAAAAATTTTGCCATATTTGCAAATATTTTTAAAAATTGCTATATTTGCAATTATTTACCTTCATGTTGCCACCCATTACAATTATCCTTTAATATATTGTTTTATTTGAAAAAAATTAAAAATATATTTTTTAAAATAAAAACCAATTGCCATTCATTCTGTTTCAAGGACGAAAAGTGTAATGAAATCCAAAAGAGACCAAAAAGTGTATTTTTTCCTTACTTTACATCCGTATCAGGAGGCTCGGTTGGTTTGAATTACCGCCCTAATCTTATTTACTGGAGAGCAAAGTGAATGGAAGTAGGTTTTAATTAGATTTGAAGGTCTCCAGTTCATGAACCGGGAATTCTTTGCTATTCTCCTGTAATTGGTGTTTTCAATATTCTAGTCCGTGGCTGGTTAGTTAGGAAATTCCAAAATTCAGCGACCAGGGTCAGGGGGCGATTTGCCCCAGGGTGGCGAGTACAGCCCTTATCCTATTGCCCGGAGGGTAAATTCCTCCCATACGGTGGTCATGGATGATGACATTGAAAATGCTAGTTTGGGGGCATACCAGGACAGGCAAAGGACTTTTCCTGACATGCGTTGTAAAGTCAACACCCCATTGGTGAGTACCTTAACCCTGCCAATCATCTGCGTTCTAATTTCAATGTATGTTGATATATTTACTGCTGGTTTGAAAAACGTGATTTTCTGACTTCTACCATGTTCTTCTATTGTATTATCTTCTTAATGACTTAACTAGCAATATTATGAAGTTATATGGTTATTTAGTTGCTTAGCATATCACAACTGAACTTGATGGACGCTATATTTGAATTACATTGTCTTGTATGATTAAGGTCTAGGTTGTTCCAGGTCACATCCTGTTGTTATTTTTTTGTATTGGATTTTACAGGGTTTAGGGTTTTAA

mRNA sequence

ATGAGACAGATTCTCAAGGCCTCCGCTGTGGGCAGCATAATGTATGCAATGCTCTGCACAAGGCCTGACATCTGTGATGATGTAGGGATAGTTAGCACATATCAATCATTAGGAAATTCCAAAATTCAGCGACCAGGGTCAGGGGGCGATTTGCCCCAGGGTGGCGAGTACAGCCCTTATCCTATTGCCCGGAGGGTAAATTCCTCCCATACGGTGGTCATGGATGATGACATTGAAAATGCTAGTTTGGGGGCATACCAGGACAGGCAAAGGACTTTTCCTGACATGCGTTGTAAAGTCAACACCCCATTGGGTTTAGGGTTTTAA

Coding sequence (CDS)

ATGAGACAGATTCTCAAGGCCTCCGCTGTGGGCAGCATAATGTATGCAATGCTCTGCACAAGGCCTGACATCTGTGATGATGTAGGGATAGTTAGCACATATCAATCATTAGGAAATTCCAAAATTCAGCGACCAGGGTCAGGGGGCGATTTGCCCCAGGGTGGCGAGTACAGCCCTTATCCTATTGCCCGGAGGGTAAATTCCTCCCATACGGTGGTCATGGATGATGACATTGAAAATGCTAGTTTGGGGGCATACCAGGACAGGCAAAGGACTTTTCCTGACATGCGTTGTAAAGTCAACACCCCATTGGGTTTAGGGTTTTAA

Protein sequence

MRQILKASAVGSIMYAMLCTRPDICDDVGIVSTYQSLGNSKIQRPGSGGDLPQGGEYSPYPIARRVNSSHTVVMDDDIENASLGAYQDRQRTFPDMRCKVNTPLGLGF
Homology
BLAST of Lag0003227.1 vs. NCBI nr
Match: KAA0059327.1 (pyrophosphate-energized membrane proton pump 2-like [Cucumis melo var. makuwa])

HSP 1 Score: 130.2 bits (326), Expect = 1.1e-26
Identity = 59/65 (90.77%), Postives = 63/65 (96.92%), Query Frame = 0

Query: 40  SKIQRPGSGGDLPQGGEYSPYPIARRVNSSHTVVMDDDIENASLGAYQDRQRTFPDMRCK 99
           SKIQRPGSGGDLPQGGEY+PYP+ARRVNSSHTV+MDDDIE+ASLG YQDRQRTFPDMR K
Sbjct: 730 SKIQRPGSGGDLPQGGEYNPYPVARRVNSSHTVLMDDDIESASLGVYQDRQRTFPDMRSK 789

Query: 100 VNTPL 105
           VNTPL
Sbjct: 790 VNTPL 794

BLAST of Lag0003227.1 vs. NCBI nr
Match: TYK04000.1 (pyrophosphate-energized membrane proton pump 2-like [Cucumis melo var. makuwa])

HSP 1 Score: 130.2 bits (326), Expect = 1.1e-26
Identity = 59/65 (90.77%), Postives = 63/65 (96.92%), Query Frame = 0

Query: 40  SKIQRPGSGGDLPQGGEYSPYPIARRVNSSHTVVMDDDIENASLGAYQDRQRTFPDMRCK 99
           SKIQRPGSGGDLPQGGEY+PYP+ARRVNSSHTV+MDDDIE+ASLG YQDRQRTFPDMR K
Sbjct: 725 SKIQRPGSGGDLPQGGEYNPYPVARRVNSSHTVLMDDDIESASLGVYQDRQRTFPDMRSK 784

Query: 100 VNTPL 105
           VNTPL
Sbjct: 785 VNTPL 789

BLAST of Lag0003227.1 vs. NCBI nr
Match: KAG7013924.1 (Pyrophosphate-energized membrane proton pump 3, partial [Cucurbita argyrosperma subsp. argyrosperma])

HSP 1 Score: 129.0 bits (323), Expect = 2.4e-26
Identity = 60/67 (89.55%), Postives = 63/67 (94.03%), Query Frame = 0

Query: 38  GNSKIQRPGSGGDLPQGGEYSPYPIARRVNSSHTVVMDDDIENASLGAYQDRQRTFPDMR 97
           G SKIQRPGSGGDLPQGGE++PYPIARRVNSSHTVVMDDDIE+ASLG YQDR RTFPDMR
Sbjct: 50  GISKIQRPGSGGDLPQGGEFNPYPIARRVNSSHTVVMDDDIESASLGVYQDRPRTFPDMR 109

Query: 98  CKVNTPL 105
            KVNTPL
Sbjct: 110 SKVNTPL 116

BLAST of Lag0003227.1 vs. NCBI nr
Match: CAB4302606.1 (unnamed protein product [Prunus armeniaca])

HSP 1 Score: 107.1 bits (266), Expect = 9.8e-20
Identity = 49/75 (65.33%), Postives = 60/75 (80.00%), Query Frame = 0

Query: 30  IVSTYQSLGNSKIQRPGSGGDLPQGGEYSPYPIARRVNSSHTVVMDDDIENASLGAYQDR 89
           I+S+     N KIQRPGSG DLPQGGEYSP+P+AR+VNSSH++VMD D+E+ +LG YQDR
Sbjct: 31  ILSSEDCWTNFKIQRPGSGADLPQGGEYSPFPVARKVNSSHSMVMDVDVEDGNLGPYQDR 90

Query: 90  QRTFPDMRCKVNTPL 105
            RTFP+MR K   PL
Sbjct: 91  PRTFPNMRGKSYNPL 105

BLAST of Lag0003227.1 vs. NCBI nr
Match: RXH67506.1 (hypothetical protein DVH24_027653 [Malus domestica])

HSP 1 Score: 104.8 bits (260), Expect = 4.9e-19
Identity = 47/64 (73.44%), Postives = 54/64 (84.38%), Query Frame = 0

Query: 41  KIQRPGSGGDLPQGGEYSPYPIARRVNSSHTVVMDDDIENASLGAYQDRQRTFPDMRCKV 100
           KIQRPGSGGDLPQGGEYSP+P+AR+VNS H+VVMD D+E  +LG YQDR RTFP+MR K 
Sbjct: 5   KIQRPGSGGDLPQGGEYSPFPVARKVNSIHSVVMDVDVEGGNLGPYQDRPRTFPNMRGKS 64

Query: 101 NTPL 105
             PL
Sbjct: 65  YNPL 68

BLAST of Lag0003227.1 vs. ExPASy Swiss-Prot
Match: Q9FWR2 (Pyrophosphate-energized membrane proton pump 3 OS=Arabidopsis thaliana OX=3702 GN=AVPL2 PE=3 SV=1)

HSP 1 Score: 46.2 bits (108), Expect = 2.7e-04
Identity = 18/33 (54.55%), Postives = 26/33 (78.79%), Query Frame = 0

Query: 72  VVMDDDIENASLGAYQDRQRTFPDMRCKVNTPL 105
           ++MD+D+E A+L +Y D+ RTFPDMR K  +PL
Sbjct: 1   MMMDEDVEQATLVSYSDKPRTFPDMRSKTYSPL 33

BLAST of Lag0003227.1 vs. ExPASy Swiss-Prot
Match: P10978 (Retrovirus-related Pol polyprotein from transposon TNT 1-94 OS=Nicotiana tabacum OX=4097 PE=2 SV=1)

HSP 1 Score: 45.4 bits (106), Expect = 4.6e-04
Identity = 20/34 (58.82%), Postives = 27/34 (79.41%), Query Frame = 0

Query: 1    MRQILKASAVGSIMYAMLCTRPDICDDVGIVSTY 35
            M ++  +SAVGS+MYAM+CTRPDI   VG+VS +
Sbjct: 1106 MAKVPYSSAVGSLMYAMVCTRPDIAHAVGVVSRF 1139

The following BLAST results are available for this feature:
Match NameE-valueIdentityDescription
KAA0059327.11.1e-2690.77pyrophosphate-energized membrane proton pump 2-like [Cucumis melo var. makuwa][more]
TYK04000.11.1e-2690.77pyrophosphate-energized membrane proton pump 2-like [Cucumis melo var. makuwa][more]
KAG7013924.12.4e-2689.55Pyrophosphate-energized membrane proton pump 3, partial [Cucurbita argyrosperma ... [more]
CAB4302606.19.8e-2065.33unnamed protein product [Prunus armeniaca][more]
RXH67506.14.9e-1973.44hypothetical protein DVH24_027653 [Malus domestica][more]
Match NameE-valueIdentityDescription
Q9FWR22.7e-0454.55Pyrophosphate-energized membrane proton pump 3 OS=Arabidopsis thaliana OX=3702 G... [more]
P109784.6e-0458.82Retrovirus-related Pol polyprotein from transposon TNT 1-94 OS=Nicotiana tabacum... [more]
Relationships

This mRNA is a part of the following gene feature(s):

Feature NameUnique NameType
Lag0003227Lag0003227gene


The following exon feature(s) are a part of this mRNA:

Feature NameUnique NameType
Lag0003227.1.exon3Lag0003227.1.exon3exon
Lag0003227.1.exon2Lag0003227.1.exon2exon
Lag0003227.1.exon1Lag0003227.1.exon1exon


The following CDS feature(s) are a part of this mRNA:

Feature NameUnique NameType
cds.Lag0003227.1cds.Lag0003227.1_3CDS
cds.Lag0003227.1cds.Lag0003227.1_2CDS
cds.Lag0003227.1cds.Lag0003227.1CDS


The following polypeptide feature(s) derives from this mRNA:

Feature NameUnique NameType
Lag0003227.1Lag0003227.1-proteinpolypeptide


GO Annotation
GO Assignments
This mRNA is annotated with the following GO terms.
Category Term Accession Term Name
biological_process GO:1902600 proton transmembrane transport
biological_process GO:0016104 triterpenoid biosynthetic process
cellular_component GO:0016021 integral component of membrane
cellular_component GO:0005811 lipid droplet
molecular_function GO:0042300 beta-amyrin synthase activity
molecular_function GO:0004427 inorganic diphosphatase activity
molecular_function GO:0000250 lanosterol synthase activity
molecular_function GO:0009678 pyrophosphate hydrolysis-driven proton transmembrane transporter activity