Cucsat.G2820.T2 (mRNA) Cucumber (B10) v3

Overview


Name	Cucsat.G2820.T2
Type	mRNA
Organism	Cucumis sativus L. var. sativus cv B10 (Cucumber (B10) v3)
Description	EPIDERMAL PATTERNING FACTOR-like protein 9
Location	ctg1041: 2943998 .. 2945699 (-)
RNA-Seq Expression	Cucsat.G2820.T2
Synteny	Cucsat.G2820.T2

Sequences

The following sequences are available for this feature:

Gene sequence (with intron)

Legend: exonpolypeptideCDS

Hold the cursor over a type above to highlight its positions in the sequence below.

TATTATTTCACACCACAAATATATATATACACACATATTTTCTTCATTATTCCCATTATTATTGTTTCCTTCAAAAACCTTCTCTTCTCTTTTTCTTTTCTCTTATCCTCTCTTTTCCAATGGCAAAAGCTGCAAAACCACTTTCAGTACTTCTTCTTCTTCTTCTTCTTCTTCTTCTTCTAACCCTATTTATTACCTCTCTTTTCTCAACAATGGCAGTGCAAGGTCCCTCTCCCTCTTATCTCTTTCTCTTTTTCTCCTCATTCATCTTCTCACCTTTTTCACTCACATGACCTTTTTCTTCTCTTTTTTTGGTTCCAGGATTCTCCACTCGAGATCGGCCAGTACCCCAATTCTCTTCTCTACAACCAACAATACTCCCACTCGAGGTTAGTCGCATTAGCCAAATAATCTCGTTTTTTTCTACCCTATATATTTCACATGAACAATACTGTTTTTCGAATTCTACCTTTAGTTTAATAATGTTATTGTTAATTTCTTTTTACTTCTACTAGTTTTGTTTTTGTGTTGGTATTGATTTAATTTTGTGTGTCATTTTCATTCAAATTTGAAATTTAAACTATAGTTTTTGGTGTAACTATCTTTTGTTTGGATTAACCTTTTTTTTTTTTTTTTTAATTTATAAAAACAACTTATAGTGATTTTAATTCATGTTTTTAAATTACCCATTGATATAGGGGCTTTTTTATGTTTCTTTTTCTAATTAATAAAAATTTATATGAACGGTTGTTTCAATCTATATTTATAAAATAAGAGTTAACCAAAATATAATGAAATAAGCATAAAAAAAATTCAAAACTAGAATCTTGAATTCACTTTTATAGTGTAATTTATTTTACCTTTTTTCTATATATATATAAAAAGAAAAGTAAAATAATATATTTTATTTTCATATATATAGAAACTTGAAAAACTTGTCTTTATAAAAGCTTTATTATATTAAGTTTCAACTTTCATTTCACAAACTACTTTAATCTCTATATACACATGCTCCAATTTTTAGATAGGAGAAAGAGAAATATATATATATATATATATATATATAATATATATATATATATATATATATATATAATATATATATATATATGTTTTAACATTTTTAGTTAACATATATTATTATTTTTGGTCACTCTCATACAATCATCATAATTGAAAGAAAATGGAGGTATGAATGATAAGAGAAAGCAAATTGAAATGAAGGAGGAGTAGTAGTAATTAAAGAGCTAATTTAGAAGTGTTATATTTATGGAGTTTAGGGTAGAAAAGAAAGGTTGAGGAAATGGAGAAGGGTGATGATTGGTTCAACAGCACCAACATGTACTTACAATGAATGTAGAGGATGCAAATACAAATGTAGAGCTGAGCAAGTTCCTGTTGAAGGCAATGACCCAATTAATAGTGCTTATCACTATAGATGTGTTTGTCATAGGTAATTTCTTCATTACTTTTATATTTCAATCCTCTTCCAAATTTTGCTCTCTGCTTTTTCTCTTAATGTCTAATCTTTTTTTTCTTTTTCTCAAATTCATTCTTTTTCTTTAAACTCTTCTTCTTCTTATTGTTATTGTAATTGTAAGTCACTTGAACTTATAAATATATATATATATATTTTTCTTTGAATGGTGTATTTTGGTTCAAGAGATATATATTCTTAGAATATATCATATTTTTAAACATT

Coding sequence (CDS)

TTATTTCACACCACAAATATATATATACACACATATTTTCTTCATTATTCCCATTATTATTGTTTCCTTCAAAAACCTTCTCTTCTCTTTTTCTTTTCTCTTATCCTCTCTTTTCCAATGGCAAAAGCTGCAAAACCACTTTCAGTACTTCTTCTTCTTCTTCTTCTTCTTCTTCTTCTAACCCTATTTATTACCTCTCTTTTCTCAACAATGGCAGTGCAAGGATTCTCCACTCGAGATCGGCCAGTACCCCAATTCTCTTCTCTACAACCAACAATACTCCCACTCGAGGGTAGAAAAGAAAGGTTGAGGAAATGGAGAAGGGTGATGATTGGTTCAACAGCACCAACATGTACTTACAATGAATGTAGAGGATGCAAATACAAATGTAGAGCTGAGCAAGTTCCTGTTGAAGGCAATGACCCAATTAATAGTGCTTATCACTATAGATGTGTTTGTCATAGGTAA

Protein sequence

LFHTTNIYIHTYFLHYSHYYCFLQKPSLLFFFSLILSFPMAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGRKERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR

Homology

[Close]

BLAST of Cucsat.G2820.T2 vs. ExPASy Swiss-Prot
Match: Q9SV72 (EPIDERMAL PATTERNING FACTOR-like protein 9 OS=Arabidopsis thaliana OX=3702 GN=EPFL9 PE=1 SV=1)

HSP 1 Score: 111.3 bits (277), Expect = 9.8e-24
Identity = 61/111 (54.95%), Postives = 70/111 (63.06%), Query Frame = 0


Query: 45  KPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGRKERLR 104
           KP  + +  LL  LLL  ++            ++R R +    SL P +  L  R     
Sbjct: 9   KPRCITIFFLLFALLLGNYVVQ----------ASRPRSIENTVSLLPQVHLLNSR----- 68

Query: 105 KWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 156
             RR MIGSTAPTCTYNECRGC+YKCRAEQVPVEGNDPINSAYHYRCVCHR
Sbjct: 69  --RRHMIGSTAPTCTYNECRGCRYKCRAEQVPVEGNDPINSAYHYRCVCHR 102

[Close]

BLAST of Cucsat.G2820.T2 vs. NCBI nr
Match: XP_031745008.1 (EPIDERMAL PATTERNING FACTOR-like protein 9 isoform X3 [Cucumis sativus])

HSP 1 Score: 229 bits (584), Expect = 7.49e-75
Identity = 116/116 (100.00%), Postives = 116/116 (100.00%), Query Frame = 0


Query: 40  MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR 99
           MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR
Sbjct: 1   MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR 60

Query: 100 KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 155
           KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR
Sbjct: 61  KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 116

[Close]

BLAST of Cucsat.G2820.T2 vs. NCBI nr
Match: XP_031745007.1 (EPIDERMAL PATTERNING FACTOR-like protein 9 isoform X2 [Cucumis sativus])

HSP 1 Score: 229 bits (584), Expect = 9.13e-75
Identity = 116/116 (100.00%), Postives = 116/116 (100.00%), Query Frame = 0


Query: 40  MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR 99
           MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR
Sbjct: 1   MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR 60

Query: 100 KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 155
           KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR
Sbjct: 61  KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 116

[Close]

BLAST of Cucsat.G2820.T2 vs. NCBI nr
Match: XP_031745006.1 (EPIDERMAL PATTERNING FACTOR-like protein 9 isoform X1 [Cucumis sativus] >KAE8646265.1 hypothetical protein Csa_016835 [Cucumis sativus])

HSP 1 Score: 229 bits (584), Expect = 1.15e-74
Identity = 116/116 (100.00%), Postives = 116/116 (100.00%), Query Frame = 0


Query: 40  MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR 99
           MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR
Sbjct: 1   MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR 60

Query: 100 KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 155
           KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR
Sbjct: 61  KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 116

[Close]

BLAST of Cucsat.G2820.T2 vs. NCBI nr
Match: XP_038899071.1 (EPIDERMAL PATTERNING FACTOR-like protein 9 [Benincasa hispida])

HSP 1 Score: 164 bits (415), Expect = 2.39e-49
Identity = 89/116 (76.72%), Postives = 95/116 (81.90%), Query Frame = 0


Query: 40  MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGR 99
           MAKAAKP+SVLLLLL      TL +T LFSTMAVQGFSTR  P  QF SLQ TI   EG+
Sbjct: 1   MAKAAKPVSVLLLLL------TLIMTFLFSTMAVQGFSTRHPP--QFFSLQATIAS-EGK 60

Query: 100 KERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 155
            E++R  RR+MIGSTAP CTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR
Sbjct: 61  NEKMRNSRRLMIGSTAPICTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 107

[Close]

BLAST of Cucsat.G2820.T2 vs. NCBI nr
Match: KAG6593029.1 (EPIDERMAL PATTERNING FACTOR-like protein 9, partial [Cucurbita argyrosperma subsp. sororia])

HSP 1 Score: 149 bits (376), Expect = 2.17e-43
Identity = 84/121 (69.42%), Postives = 92/121 (76.03%), Query Frame = 0


Query: 40  MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQG-FSTRD----RPVPQFSSLQPTIL 99
           MAKAAKPL      LLLLLLL L + SL STMA++G FST D    +P+PQF S+Q TI 
Sbjct: 1   MAKAAKPL------LLLLLLLALILCSLCSTMALKGRFSTEDELFAKPLPQFRSVQTTIA 60

Query: 100 PLEGRKERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCH 155
           P       +R  RR+MIGSTAP CTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCH
Sbjct: 61  P------EMRNSRRLMIGSTAPICTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCH 109

[Close]

BLAST of Cucsat.G2820.T2 vs. ExPASy TrEMBL
Match: A0A0A0KA17 (EPIDERMAL PATTERNING FACTOR-like protein 9 OS=Cucumis sativus OX=3659 GN=Csa_7G368090 PE=4 SV=1)

HSP 1 Score: 187 bits (475), Expect = 4.28e-59
Identity = 85/85 (100.00%), Postives = 85/85 (100.00%), Query Frame = 0


Query: 71  MAVQGFSTRDRPVPQFSSLQPTILPLEGRKERLRKWRRVMIGSTAPTCTYNECRGCKYKC 130
           MAVQGFSTRDRPVPQFSSLQPTILPLEGRKERLRKWRRVMIGSTAPTCTYNECRGCKYKC
Sbjct: 1   MAVQGFSTRDRPVPQFSSLQPTILPLEGRKERLRKWRRVMIGSTAPTCTYNECRGCKYKC 60

Query: 131 RAEQVPVEGNDPINSAYHYRCVCHR 155
           RAEQVPVEGNDPINSAYHYRCVCHR
Sbjct: 61  RAEQVPVEGNDPINSAYHYRCVCHR 85

[Close]

BLAST of Cucsat.G2820.T2 vs. ExPASy TrEMBL
Match: A0A6J1H996 (EPIDERMAL PATTERNING FACTOR-like protein 9 OS=Cucurbita moschata OX=3662 GN=LOC111460784 PE=4 SV=1)

HSP 1 Score: 148 bits (373), Expect = 3.10e-43
Identity = 84/121 (69.42%), Postives = 92/121 (76.03%), Query Frame = 0


Query: 40  MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQG-FSTRD----RPVPQFSSLQPTIL 99
           MAKAAKPL     L LLLLLL L + SL STMA++G FST D    +P+PQF S+Q TI 
Sbjct: 1   MAKAAKPL-----LRLLLLLLALILCSLCSTMALKGRFSTEDELFAKPLPQFRSVQTTIA 60

Query: 100 PLEGRKERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCH 155
           P       +R  RR+MIGSTAP CTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCH
Sbjct: 61  P------EMRNSRRLMIGSTAPICTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCH 110

[Close]

BLAST of Cucsat.G2820.T2 vs. ExPASy TrEMBL
Match: A0A6J1KTW3 (EPIDERMAL PATTERNING FACTOR-like protein 9 OS=Cucurbita maxima OX=3661 GN=LOC111497591 PE=4 SV=1)

HSP 1 Score: 145 bits (365), Expect = 4.80e-42
Identity = 82/121 (67.77%), Postives = 90/121 (74.38%), Query Frame = 0


Query: 40  MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQG-FSTRD----RPVPQFSSLQPTIL 99
           MAKAAKPL       LLLLLL L + SL STMA++G FST D    + +PQF S+Q TI 
Sbjct: 1   MAKAAKPL-------LLLLLLALILCSLCSTMALKGRFSTEDDVFAKSLPQFRSVQTTIA 60

Query: 100 PLEGRKERLRKWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCH 155
           P       +R  RR+MIGSTAP CTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCH
Sbjct: 61  P------EMRNSRRLMIGSTAPICTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCH 108

[Close]

BLAST of Cucsat.G2820.T2 vs. ExPASy TrEMBL
Match: A0A6J1DR94 (uncharacterized protein LOC111023601 OS=Momordica charantia OX=3673 GN=LOC111023601 PE=4 SV=1)

HSP 1 Score: 137 bits (344), Expect = 8.93e-39
Identity = 77/125 (61.60%), Postives = 90/125 (72.00%), Query Frame = 0


Query: 40  MAKAAKPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPV-------PQFSSLQPT 99
           MAKAAKP SVLLLLL           +L   +AVQGF+T +  +       PQFSS+  +
Sbjct: 1   MAKAAKPASVLLLLL-----------TLIIVVAVQGFTTGEDELLLAKALPPQFSSV--S 60

Query: 100 ILPLEGRKERLR--KWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYR 155
            +  +G+ + LR  + RR+MIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYR
Sbjct: 61  TMANQGKNKMLRNSRTRRLMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYR 112

[Close]

BLAST of Cucsat.G2820.T2 vs. ExPASy TrEMBL
Match: A0A2R6P6P2 (Stomagen like OS=Actinidia chinensis var. chinensis OX=1590841 GN=CEY00_Acc31935 PE=4 SV=1)

HSP 1 Score: 124 bits (311), Expect = 8.56e-34
Identity = 66/104 (63.46%), Postives = 72/104 (69.23%), Query Frame = 0


Query: 59  LLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQ-----PTILPLEGRKERL--RKWRRVMI 118
           LL LF T +F+   +QG S     VP   SL+      T LP  G  E    R  RR+MI
Sbjct: 9   LLLLFFTIIFAAHVIQGSSRTPFSVPSIESLRRSSASETELPQPGGFEPWVSRNSRRLMI 68

Query: 119 GSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 155
           GSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHY+CVCHR
Sbjct: 69  GSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYKCVCHR 112

[Close]

BLAST of Cucsat.G2820.T2 vs. TAIR 10
Match: AT4G12970.1 (stomagen )

HSP 1 Score: 111.3 bits (277), Expect = 7.0e-25
Identity = 61/111 (54.95%), Postives = 70/111 (63.06%), Query Frame = 0


Query: 45  KPLSVLLLLLLLLLLLTLFITSLFSTMAVQGFSTRDRPVPQFSSLQPTILPLEGRKERLR 104
           KP  + +  LL  LLL  ++            ++R R +    SL P +  L  R     
Sbjct: 9   KPRCITIFFLLFALLLGNYVVQ----------ASRPRSIENTVSLLPQVHLLNSR----- 68

Query: 105 KWRRVMIGSTAPTCTYNECRGCKYKCRAEQVPVEGNDPINSAYHYRCVCHR 156
             RR MIGSTAPTCTYNECRGC+YKCRAEQVPVEGNDPINSAYHYRCVCHR
Sbjct: 69  --RRHMIGSTAPTCTYNECRGCRYKCRAEQVPVEGNDPINSAYHYRCVCHR 102

The following BLAST results are available for this feature:

BLAST of Cucsat.G2820.T2 vs. ExPASy Swiss-Prot


Match Name	E-value	Identity	Description
Q9SV72	9.8e-24	54.95	EPIDERMAL PATTERNING FACTOR-like protein 9 OS=Arabidopsis thaliana OX=3702 GN=EP...	[more]

BLAST of Cucsat.G2820.T2 vs. NCBI nr


Match Name	E-value	Identity	Description
XP_031745008.1	7.49e-75	100.00	EPIDERMAL PATTERNING FACTOR-like protein 9 isoform X3 [Cucumis sativus]	[more]
XP_031745007.1	9.13e-75	100.00	EPIDERMAL PATTERNING FACTOR-like protein 9 isoform X2 [Cucumis sativus]	[more]
XP_031745006.1	1.15e-74	100.00	EPIDERMAL PATTERNING FACTOR-like protein 9 isoform X1 [Cucumis sativus] >KAE8646...	[more]
XP_038899071.1	2.39e-49	76.72	EPIDERMAL PATTERNING FACTOR-like protein 9 [Benincasa hispida]	[more]
KAG6593029.1	2.17e-43	69.42	EPIDERMAL PATTERNING FACTOR-like protein 9, partial [Cucurbita argyrosperma subs...	[more]

BLAST of Cucsat.G2820.T2 vs. ExPASy TrEMBL


Match Name	E-value	Identity	Description
A0A0A0KA17	4.28e-59	100.00	EPIDERMAL PATTERNING FACTOR-like protein 9 OS=Cucumis sativus OX=3659 GN=Csa_7G3...	[more]
A0A6J1H996	3.10e-43	69.42	EPIDERMAL PATTERNING FACTOR-like protein 9 OS=Cucurbita moschata OX=3662 GN=LOC1...	[more]
A0A6J1KTW3	4.80e-42	67.77	EPIDERMAL PATTERNING FACTOR-like protein 9 OS=Cucurbita maxima OX=3661 GN=LOC111...	[more]
A0A6J1DR94	8.93e-39	61.60	uncharacterized protein LOC111023601 OS=Momordica charantia OX=3673 GN=LOC111023...	[more]
A0A2R6P6P2	8.56e-34	63.46	Stomagen like OS=Actinidia chinensis var. chinensis OX=1590841 GN=CEY00_Acc31935...	[more]

BLAST of Cucsat.G2820.T2 vs. TAIR 10


Match Name	E-value	Identity	Description
AT4G12970.1	7.0e-25	54.95	stomagen	[more]

InterPro

Analysis Name: InterPro Annotations of Cucumber (B10) v3
Date Performed: 2021-10-25


IPR Term	IPR Description	Source	Source Term	Source Description	Alignment
IPR038572	Stomagen, C-terminal domain superfamily	GENE3D	2.20.25.390	Stomagen	coord: 111..155 e-value: 4.3E-31 score: 108.5
IPR044858	Stomagen, C-terminal domain	PFAM	PF16851	Stomagen	coord: 107..154 e-value: 5.4E-33 score: 112.8
IPR031753	Stomagen	PANTHER	PTHR37239	EPIDERMAL PATTERNING FACTOR-LIKE PROTEIN 9	coord: 58..155

Relationships

This mRNA is a part of the following gene feature(s):


Feature Name	Unique Name	Type
Cucsat.G2820	Cucsat.G2820	gene

The following exon feature(s) are a part of this mRNA:


Feature Name	Unique Name	Type
Cucsat.G2820.T2.E1	Cucsat.G2820.T2.E1	exon
Cucsat.G2820.T2.E2	Cucsat.G2820.T2.E2	exon
Cucsat.G2820.T2.E3	Cucsat.G2820.T2.E3	exon

The following CDS feature(s) are a part of this mRNA:


Feature Name	Unique Name	Type
Cucsat.G2820.T2.C3	Cucsat.G2820.T2.C3	CDS
Cucsat.G2820.T2.C2	Cucsat.G2820.T2.C2	CDS
Cucsat.G2820.T2.C1	Cucsat.G2820.T2.C1	CDS

The following polypeptide feature(s) derives from this mRNA:


Feature Name	Unique Name	Type
Cucsat.G2820.T2	Cucsat.G2820.T2-protein	polypeptide

GO Annotation

GO Assignments

This mRNA is annotated with the following GO terms.

Category	Term Accession	Term Name
biological_process	GO:2000123	positive regulation of stomatal complex development