Lag0027498 (gene) Sponge gourd (AG‐4) v1

Overview
NameLag0027498
Typegene
OrganismLuffa acutangula (Sponge gourd (AG‐4) v1)
Description30S ribosomal protein S21, chloroplastic-like
Locationchr8: 1450674 .. 1452291 (+)
RNA-Seq ExpressionLag0027498
SyntenyLag0027498
Sequences
The following sequences are available for this feature:

Gene sequence (with intron)

Legend: exonCDSpolypeptide
Hold the cursor over a type above to highlight its positions in the sequence below.
ATGGCTTCTCTCTCCACCTTCTTCTCCTTCCTCTTACCCTCAAAACCTCCGCCCCCCAAACCCCATCAGCCGCTCATTGCCTCTCCTTCTCCGGCAGCCCCTCAGAACATTCAATTCCAATCCCTCAAATCCAGAGATGGATCGGTTGCCCTAAACCCCCACGAGGGTTCGAACCATTCCGAGCCCTTGTCGGCCGAATTGGCCTCCGTTATCTGTCCCTCGCTGGCCTTCGCCAACACTATGTTCTTCCGATCGGCGTATAATGTGCAGGTGATCGTGGATGATAACGAGCCGGAGGAGCGGCTTCTGAACAGATTCCGGCGGGAGGTTATGAGGGCCGGCGTGATTCAGGAGTGTAAGAGGCGGAGATTCTTTGAGAATACTCAGGATGTCAGGAAGCGCAAGTCTCGCGAGGCTGCGAAACGCAACCGCCGGAGGTCTGTTTCTGAAATCTCTTGTGGTTTGGTTTCTTTTGGGATTTTTGCTCTTGGTTTCTGAAACTCTCTGATTGTTTGTTTGTTGGGATTTTCATTGATTCTTGGAGCTGTTTGGGATTTGTGTTCTGTTTGAGTGTTGAATTTGATTGCTAAGAAGGTAGTAGGAGTCTTATCCAGCGGTTTGGAGAGATTTATCTATAGTTTGGAGGGGACAAGATTTGAACTTTTGACCTCATGGTCGAGGATACATGCGAATTACCATTGAGCTATGCTCACTTTGACAAGATTTATATAGAGTTACTCCTGACTGGACTAATGTTGTGTATTTTAAAAGTTAAACTATAATAATGTAATAAGTTCAAGCCATGAAGGTGTTACCTATCTAAGATTTAATATCTTACAAGTAAATGTAGTAGGGTCAGACAGTTGTCCTGTAAGAATAATCGAGGTGCACAAACTTGGCCTGGACACTCACGCTGTAGTAATGTAGAAGAAAAGAGAAAATAGGCACATTCCCTTGGACATGTTTTACCTTTACTCTTTCGAGCAACTTAGGATATGGAGTTCGAGTTAAGAGAAAGTAGAGAGGGAAAAAAGCGTATCTAAGGGGACTTGAGAGAAAATGGAGAGAAAAACCCTGTGGATTCACACGTTCACATCTCGAAGGTTGATGGTTCAATTTTCCACCTAATACAATGAGGATGTGGTTTTTCAATTGTTCTTCTACTTGTTGTAATCTTTGGTGTTTGTAGATAGCCAAACTTTTCTTTCTTTTTACTTTGCTTTTTCTGTTTTCGTTCCTATTGTTTATTGTGTTTGTAGATAGAGAATTTCAGATTTTGCAGAACCTTCAAATTCCTAACATCTTTATCTTGAAACCAAATATCACTTTGAAACTTCTACTCAATTTGGAAGATGGCTGTAGAGTGTCTTGTTAGAGATTTTAATGCACAATTAAGTGAATTCTTGAGCCCCCATTGGCTAATTGCAGTCTTCTGTCAAAAGAAAGTGGATGGAATTTATGTGCTAAGTTGTTTGATGAAACTGAGTTATGATTTTGGTTGATGAACAGGCGCCCACAGCCAAGATTTACACCACAGAACAAGCAGGATGTGCCTGCAACTAAGCAGGAAGGTGATGATGACAACTGGGATTTACCTGAAGAAGATGTTCCTTATTAA

mRNA sequence

ATGGCTTCTCTCTCCACCTTCTTCTCCTTCCTCTTACCCTCAAAACCTCCGCCCCCCAAACCCCATCAGCCGCTCATTGCCTCTCCTTCTCCGGCAGCCCCTCAGAACATTCAATTCCAATCCCTCAAATCCAGAGATGGATCGGTTGCCCTAAACCCCCACGAGGGTTCGAACCATTCCGAGCCCTTGTCGGCCGAATTGGCCTCCGTTATCTGTCCCTCGCTGGCCTTCGCCAACACTATGTTCTTCCGATCGGCGTATAATGTGCAGGTGATCGTGGATGATAACGAGCCGGAGGAGCGGCTTCTGAACAGATTCCGGCGGGAGGTTATGAGGGCCGGCGTGATTCAGGAGTGTAAGAGGCGGAGATTCTTTGAGAATACTCAGGATGTCAGGAAGCGCAAGTCTCGCGAGGCTGCGAAACGCAACCGCCGGAGGCGCCCACAGCCAAGATTTACACCACAGAACAAGCAGGATGTGCCTGCAACTAAGCAGGAAGGTGATGATGACAACTGGGATTTACCTGAAGAAGATGTTCCTTATTAA

Coding sequence (CDS)

ATGGCTTCTCTCTCCACCTTCTTCTCCTTCCTCTTACCCTCAAAACCTCCGCCCCCCAAACCCCATCAGCCGCTCATTGCCTCTCCTTCTCCGGCAGCCCCTCAGAACATTCAATTCCAATCCCTCAAATCCAGAGATGGATCGGTTGCCCTAAACCCCCACGAGGGTTCGAACCATTCCGAGCCCTTGTCGGCCGAATTGGCCTCCGTTATCTGTCCCTCGCTGGCCTTCGCCAACACTATGTTCTTCCGATCGGCGTATAATGTGCAGGTGATCGTGGATGATAACGAGCCGGAGGAGCGGCTTCTGAACAGATTCCGGCGGGAGGTTATGAGGGCCGGCGTGATTCAGGAGTGTAAGAGGCGGAGATTCTTTGAGAATACTCAGGATGTCAGGAAGCGCAAGTCTCGCGAGGCTGCGAAACGCAACCGCCGGAGGCGCCCACAGCCAAGATTTACACCACAGAACAAGCAGGATGTGCCTGCAACTAAGCAGGAAGGTGATGATGACAACTGGGATTTACCTGAAGAAGATGTTCCTTATTAA

Protein sequence

MASLSTFFSFLLPSKPPPPKPHQPLIASPSPAAPQNIQFQSLKSRDGSVALNPHEGSNHSEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQECKRRRFFENTQDVRKRKSREAAKRNRRRRPQPRFTPQNKQDVPATKQEGDDDNWDLPEEDVPY
Homology
[Close]
BLAST of Lag0027498 vs. NCBI nr
Match: XP_008450090.2 (PREDICTED: uncharacterized protein LOC103491780, partial [Cucumis melo])

HSP 1 Score: 308.9 bits (790), Expect = 2.9e-80
Identity = 165/183 (90.16%), Postives = 170/183 (92.90%), Query Frame = 0 

Query: 1   MASLSTFFSFLLPSKPPPPKPHQPLIA-SPSPAAPQNIQFQSLKSRDGSVALNPHEGSNH 60
           MASLS+FFSFLLPSKPPPPKPHQPLIA SPSP      QFQ+LKSRDGSVAL P +G NH
Sbjct: 49  MASLSSFFSFLLPSKPPPPKPHQPLIASSPSPPNHAAAQFQTLKSRDGSVALKPQQGPNH 108

Query: 61  SEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQEC 120
           SEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQEC
Sbjct: 109 SEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQEC 168

Query: 121 KRRRFFENTQDVRKRKSREAAKRNRRRRPQPRFTPQNKQDVPATKQEGDDDNWDLPEE-D 180
           KRRRFFENTQDVRKRK+REAAKRNRRRRPQ RFTPQNKQDVPATKQE DDDNWDLPEE D
Sbjct: 169 KRRRFFENTQDVRKRKTREAAKRNRRRRPQARFTPQNKQDVPATKQEADDDNWDLPEEAD 228

Query: 181 VPY 182
           +PY
Sbjct: 229 IPY 231

[Close]
BLAST of Lag0027498 vs. NCBI nr
Match: KAA0042889.1 (hypothetical protein E6C27_scaffold44G004060 [Cucumis melo var. makuwa] >TYK05292.1 hypothetical protein E5676_scaffold108G001120 [Cucumis melo var. makuwa])

HSP 1 Score: 308.9 bits (790), Expect = 2.9e-80
Identity = 165/183 (90.16%), Postives = 170/183 (92.90%), Query Frame = 0 

Query: 1   MASLSTFFSFLLPSKPPPPKPHQPLIA-SPSPAAPQNIQFQSLKSRDGSVALNPHEGSNH 60
           MASLS+FFSFLLPSKPPPPKPHQPLIA SPSP      QFQ+LKSRDGSVAL P +G NH
Sbjct: 1   MASLSSFFSFLLPSKPPPPKPHQPLIASSPSPPNHAAAQFQTLKSRDGSVALKPQQGPNH 60

Query: 61  SEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQEC 120
           SEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQEC
Sbjct: 61  SEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQEC 120

Query: 121 KRRRFFENTQDVRKRKSREAAKRNRRRRPQPRFTPQNKQDVPATKQEGDDDNWDLP-EED 180
           KRRRFFENTQDVRKRK+REAAKRNRRRRPQ RFTPQNKQDVPATKQE DDDNWDLP EED
Sbjct: 121 KRRRFFENTQDVRKRKTREAAKRNRRRRPQARFTPQNKQDVPATKQEADDDNWDLPEEED 180

Query: 181 VPY 182
           +PY
Sbjct: 181 IPY 183

[Close]
BLAST of Lag0027498 vs. NCBI nr
Match: AAL35983.1 (ribosome-like protein, partial [Cucumis sativus] >4V61_AU Chain AU, Ribosomal Protein S21 [Spinacia oleracea])

HSP 1 Score: 305.1 bits (780), Expect = 4.1e-79
Identity = 162/182 (89.01%), Postives = 170/182 (93.41%), Query Frame = 0 

Query: 1   MASLSTFFSFLLPSKPPPPKPHQPLIA-SPSPAAPQNIQFQSLKSRDGSVALNPHEGSNH 60
           MASLS+FFSFLLPSKPPPPKPHQP IA SPSP+     QFQ+LKSRDGSVALNP +G NH
Sbjct: 8   MASLSSFFSFLLPSKPPPPKPHQPFIASSPSPSNHAAPQFQTLKSRDGSVALNPQQGPNH 67

Query: 61  SEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQEC 120
           SEPLSAELASVICPSLA+ANTMFFRSAYNVQV+VDDNEPEERLLNRFRREVMRAGVIQEC
Sbjct: 68  SEPLSAELASVICPSLAYANTMFFRSAYNVQVVVDDNEPEERLLNRFRREVMRAGVIQEC 127

Query: 121 KRRRFFENTQDVRKRKSREAAKRNRRRRPQPRFTPQNKQDVPATKQEGDDDNWDLPE-ED 180
           KRRRFFENTQDVRKRK+REAAKRNRRRRPQ RFTPQNKQDVPATKQE DDDNWDLPE ED
Sbjct: 128 KRRRFFENTQDVRKRKTREAAKRNRRRRPQARFTPQNKQDVPATKQEADDDNWDLPEDED 187

[Close]
BLAST of Lag0027498 vs. NCBI nr
Match: XP_004145671.1 (30S ribosomal protein S21, chloroplastic [Cucumis sativus] >KGN58186.1 hypothetical protein Csa_017561 [Cucumis sativus])

HSP 1 Score: 305.1 bits (780), Expect = 4.1e-79
Identity = 162/182 (89.01%), Postives = 170/182 (93.41%), Query Frame = 0 

Query: 1   MASLSTFFSFLLPSKPPPPKPHQPLIA-SPSPAAPQNIQFQSLKSRDGSVALNPHEGSNH 60
           MASLS+FFSFLLPSKPPPPKPHQP IA SPSP+     QFQ+LKSRDGSVALNP +G NH
Sbjct: 1   MASLSSFFSFLLPSKPPPPKPHQPFIASSPSPSNHAAPQFQTLKSRDGSVALNPQQGPNH 60

Query: 61  SEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQEC 120
           SEPLSAELASVICPSLA+ANTMFFRSAYNVQV+VDDNEPEERLLNRFRREVMRAGVIQEC
Sbjct: 61  SEPLSAELASVICPSLAYANTMFFRSAYNVQVVVDDNEPEERLLNRFRREVMRAGVIQEC 120

Query: 121 KRRRFFENTQDVRKRKSREAAKRNRRRRPQPRFTPQNKQDVPATKQEGDDDNWDLPE-ED 180
           KRRRFFENTQDVRKRK+REAAKRNRRRRPQ RFTPQNKQDVPATKQE DDDNWDLPE ED
Sbjct: 121 KRRRFFENTQDVRKRKTREAAKRNRRRRPQARFTPQNKQDVPATKQEADDDNWDLPEDED 180

[Close]
BLAST of Lag0027498 vs. NCBI nr
Match: KAG6581093.1 (30S ribosomal protein S21, chloroplastic, partial [Cucurbita argyrosperma subsp. sororia])

HSP 1 Score: 303.9 bits (777), Expect = 9.2e-79
Identity = 159/181 (87.85%), Postives = 167/181 (92.27%), Query Frame = 0 

Query: 1   MASLSTFFSFLLPSKPPPPKPHQPLIASPSPAAPQNIQFQSLKSRDGSVALNPHEGSNHS 60
           MASLS+FFSFLLPSKPPPPKP QPLIASPSPA     QFQ+LKSRDGSVAL+P + S HS
Sbjct: 1   MASLSSFFSFLLPSKPPPPKPLQPLIASPSPANQAAAQFQTLKSRDGSVALSPQQASKHS 60

Query: 61  EPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQECK 120
           EPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQECK
Sbjct: 61  EPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQECK 120

Query: 121 RRRFFENTQDVRKRKSREAAKRNRRRRPQPRFTPQNKQDVPATKQEGDDDNWDLPEEDVP 180
           RR+FFEN  DV+KRKSREAAKRNRRRRPQ RFTPQ+KQDVPATKQE DDDNWDLPEE +P
Sbjct: 121 RRKFFENPHDVKKRKSREAAKRNRRRRPQQRFTPQDKQDVPATKQEADDDNWDLPEEGIP 180

Query: 181 Y 182
           Y
Sbjct: 181 Y 181

[Close]
BLAST of Lag0027498 vs. ExPASy Swiss-Prot
Match: P82024 (30S ribosomal protein S21, chloroplastic OS=Spinacia oleracea OX=3562 GN=rps21 PE=1 SV=2)

HSP 1 Score: 152.9 bits (385), Expect = 3.4e-36
Identity = 94/186 (50.54%), Postives = 118/186 (63.44%), Query Frame = 0 

Query: 2   ASLSTFFSFLLPSKPPPPKPHQPLIASPSPAAPQNIQFQSLKSRDGSVALNPHEGSN--- 61
           +SL  F S L PS    P    P   +P+          SL S+  S++++  +  N   
Sbjct: 5   SSLLNFLSPLFPSNTSLPPSSNPKFPNPN----------SLSSQQNSISISSKKHENAAI 64

Query: 62  -HSEPLSAELASVICPSLAFANTMFFRSAYNVQVIVDDNEPEERLLNRFRREVMRAGVIQ 121
              E    +L +V+CPSLAF+NT++FRSAYNVQV+VDDNE EERLLNRFRREVMRAGVIQ
Sbjct: 65  AKKEEYPGDLMAVVCPSLAFSNTLYFRSAYNVQVLVDDNENEERLLNRFRREVMRAGVIQ 124

Query: 122 ECKRRRFFENTQDVRKRKSREAAKRNRRRRPQP--RFTPQNKQDVPATKQEGDDDNWDLP 181
           ECKRRR+FEN Q+ +KRK REAAKRN RRR  P     P  ++     K+E D DNWD+P
Sbjct: 125 ECKRRRYFENKQEEKKRKHREAAKRNSRRRRGPFRGPFPGKEEATKVDKKEDDGDNWDMP 180

[Close]
BLAST of Lag0027498 vs. ExPASy Swiss-Prot
Match: P48949 (30S ribosomal protein S21 OS=Synechocystis sp. (strain PCC 6803 / Kazusa) OX=1111708 GN=rpsU PE=3 SV=1)

HSP 1 Score: 47.4 bits (111), Expect = 2.0e-04
Identity = 27/58 (46.55%), Postives = 38/58 (65.52%), Query Frame = 0 

Query: 90  QVIVDDNEPEERLLNRFRREVMRAGVIQECKRRRFFENTQDVRKRKSREAAKRNRRRR 148
           QV+V  NEP E  L RF+R+V +AG+  + K+ +FFE  Q+  KRK  EA +R +R R
Sbjct: 3   QVVVGQNEPIESALRRFKRQVAKAGIYTDFKKHQFFETPQEKHKRK--EATRRRQRSR 58

[Close]
BLAST of Lag0027498 vs. ExPASy Swiss-Prot
Match: B8GPU0 (30S ribosomal protein S21 OS=Thioalkalivibrio sulfidiphilus (strain HL-EbGR7) OX=396588 GN=rpsU PE=3 SV=1)

HSP 1 Score: 46.6 bits (109), Expect = 3.5e-04
Identity = 26/56 (46.43%), Postives = 36/56 (64.29%), Query Frame = 0 

Query: 91  VIVDDNEPEERLLNRFRREVMRAGVIQECKRRRFFENTQDVRKRKSREAAKRNRRR 147
           V V +NEP E  L RF+R   +AGV+ E +RR F+E   ++RKRK+  A KR  +R
Sbjct: 4   VRVKENEPFEVALRRFKRTCEKAGVLTEVRRREFYEKPTEIRKRKAAAAVKRQAKR 59

[Close]
BLAST of Lag0027498 vs. ExPASy Swiss-Prot
Match: A1WW12 (30S ribosomal protein S21 OS=Halorhodospira halophila (strain DSM 244 / SL1) OX=349124 GN=rpsU PE=3 SV=1)

HSP 1 Score: 45.4 bits (106), Expect = 7.7e-04
Identity = 26/56 (46.43%), Postives = 35/56 (62.50%), Query Frame = 0 

Query: 91  VIVDDNEPEERLLNRFRREVMRAGVIQECKRRRFFENTQDVRKRKSREAAKRNRRR 147
           V V +NEP E  L RF+R   +AGV+ E +RR ++E    VRKRK   A KR+ +R
Sbjct: 4   VKVRENEPFEVALRRFKRSCEKAGVLSEIRRREYYEKPTQVRKRKQAAAVKRHMKR 59

[Close]
BLAST of Lag0027498 vs. ExPASy Swiss-Prot
Match: B4U8H0 (30S ribosomal protein S21 OS=Hydrogenobaculum sp. (strain Y04AAS1) OX=380749 GN=rpsU PE=3 SV=1)

HSP 1 Score: 45.4 bits (106), Expect = 7.7e-04
Identity = 24/52 (46.15%), Postives = 33/52 (63.46%), Query Frame = 0 

Query: 91  VIVDDNEPEERLLNRFRREVMRAGVIQECKRRRFFENTQDVRKRKSREAAKR 143
           VIV D+E  E+ + RF++ V R G++ E KRR F+E     +KRK R A KR
Sbjct: 4   VIVGDSESFEKAMKRFKKVVEREGILSELKRREFYEKPSQKKKRKERAARKR 55

The following BLAST results are available for this feature:
BLAST of Lag0027498 vs. NCBI nr
Match NameE-valueIdentityDescription
XP_008450090.22.9e-8090.16PREDICTED: uncharacterized protein LOC103491780, partial [Cucumis melo][more]
KAA0042889.12.9e-8090.16hypothetical protein E6C27_scaffold44G004060 [Cucumis melo var. makuwa] >TYK0529... [more]
AAL35983.14.1e-7989.01ribosome-like protein, partial [Cucumis sativus] >4V61_AU Chain AU, Ribosomal Pr... [more]
XP_004145671.14.1e-7989.0130S ribosomal protein S21, chloroplastic [Cucumis sativus] >KGN58186.1 hypotheti... [more]
KAG6581093.19.2e-7987.8530S ribosomal protein S21, chloroplastic, partial [Cucurbita argyrosperma subsp.... [more]
BLAST of Lag0027498 vs. ExPASy Swiss-Prot
Match NameE-valueIdentityDescription
P820243.4e-3650.5430S ribosomal protein S21, chloroplastic OS=Spinacia oleracea OX=3562 GN=rps21 P... [more]
P489492.0e-0446.5530S ribosomal protein S21 OS=Synechocystis sp. (strain PCC 6803 / Kazusa) OX=111... [more]
B8GPU03.5e-0446.4330S ribosomal protein S21 OS=Thioalkalivibrio sulfidiphilus (strain HL-EbGR7) OX... [more]
A1WW127.7e-0446.4330S ribosomal protein S21 OS=Halorhodospira halophila (strain DSM 244 / SL1) OX=... [more]
B4U8H07.7e-0446.1530S ribosomal protein S21 OS=Hydrogenobaculum sp. (strain Y04AAS1) OX=380749 GN=... [more]
InterPro
Analysis Name: InterPro Annotations of Sponge gourd (AG-4) v1
Date Performed: 2022-08-01
IPR TermIPR DescriptionSourceSource TermSource DescriptionAlignment
IPR001911Ribosomal protein S21PRINTSPR00976RIBOSOMALS21coord: 92..100
score: 42.59
coord: 112..122
score: 38.64
coord: 100..109
score: 43.33
coord: 124..134
score: 37.88
IPR001911Ribosomal protein S21TIGRFAMTIGR00030TIGR00030coord: 89..146
e-value: 1.7E-13
score: 48.1
IPR001911Ribosomal protein S21PFAMPF01165Ribosomal_S21coord: 90..144
e-value: 2.6E-19
score: 68.7
IPR001911Ribosomal protein S21PANTHERPTHR21109MITOCHONDRIAL 28S RIBOSOMAL PROTEIN S21coord: 3..181
IPR001911Ribosomal protein S21HAMAPMF_00358Ribosomal_S21coord: 88..144
score: 14.053146
IPR038380Ribosomal protein S21 superfamilyGENE3D1.20.5.1150Ribosomal protein S8coord: 103..141
e-value: 1.4E-12
score: 48.8
NoneNo IPR availableMOBIDB_LITEmobidb-litedisorder_predictioncoord: 129..181
NoneNo IPR availableMOBIDB_LITEmobidb-litedisorder_predictioncoord: 137..151
NoneNo IPR availableMOBIDB_LITEmobidb-litedisorder_predictioncoord: 13..28
NoneNo IPR availableMOBIDB_LITEmobidb-litedisorder_predictioncoord: 13..33
NoneNo IPR availablePANTHERPTHR21109:SF17PROTEIN S21 FAMILY PROTEIN, PUTATIVE-RELATEDcoord: 3..181

Relationships

The following mRNA feature(s) are a part of this gene:

Feature NameUnique NameType
Lag0027498.1Lag0027498.1mRNA


GO Annotation
GO Assignments
This gene is annotated with the following GO terms.
Category Term Accession Term Name
biological_process GO:0006412 translation
cellular_component GO:0005840 ribosome
molecular_function GO:0003735 structural constituent of ribosome