CcUC04G069160 (gene) Watermelon (PI 537277) v1

Overview


Name	CcUC04G069160
Type	gene
Organism	Citrullus colocynthis (Watermelon (PI 537277) v1)
Description	cysteine proteinase inhibitor 6-like
Location	CicolChr04: 25802940 .. 25803302 (-)
RNA-Seq Expression	CcUC04G069160
Synteny	CcUC04G069160

Sequences

The following sequences are available for this feature:

Gene sequence (with intron)

Legend: exonCDSpolypeptide

Hold the cursor over a type above to highlight its positions in the sequence below.

ATGGAATTCGAGATAGTCGCCCCGGGACAAAACGATGCCTTGCTCCAAAACCGGTGCGAGCCAGGTGGTTGGGAACCGATCGAGAACATCCGCGACCCGTACGTGCAAGAGATGGGAAGGTTTGCAGTGATGGAACACACACACAAGCTAGTAGGAGCATCACTGAGGTTCATTCGTGTGGTGAGTGGTGAGACTAGGCCGATGAGCGAAGGCAAAGAATACAGGCTTGTGGTGGAAGTGGCAGAACAGATAATAACTAGTCCTCCAATGCTATCAGTTTCTATCAAGTTTTACTTGGCAACTGTTCTGGAGAATCCATTAGATCGATCTTGGAAACTCAAAGCTTTTGTGCCTTGTAATTAG

mRNA sequence

ATGGAATTCGAGATAGTCGCCCCGGGACAAAACGATGCCTTGCTCCAAAACCGGTGCGAGCCAGGTGGTTGGGAACCGATCGAGAACATCCGCGACCCGTACGTGCAAGAGATGGGAAGGTTTGCAGTGATGGAACACACACACAAGCTAGTAGGAGCATCACTGAGGTTCATTCGTGTGGTGAGTGGTGAGACTAGGCCGATGAGCGAAGGCAAAGAATACAGGCTTGTGGTGGAAGTGGCAGAACAGATAATAACTAGTCCTCCAATGCTATCAGTTTCTATCAAGTTTTACTTGGCAACTGTTCTGGAGAATCCATTAGATCGATCTTGGAAACTCAAAGCTTTTGTGCCTTGTAATTAG

Coding sequence (CDS)

ATGGAATTCGAGATAGTCGCCCCGGGACAAAACGATGCCTTGCTCCAAAACCGGTGCGAGCCAGGTGGTTGGGAACCGATCGAGAACATCCGCGACCCGTACGTGCAAGAGATGGGAAGGTTTGCAGTGATGGAACACACACACAAGCTAGTAGGAGCATCACTGAGGTTCATTCGTGTGGTGAGTGGTGAGACTAGGCCGATGAGCGAAGGCAAAGAATACAGGCTTGTGGTGGAAGTGGCAGAACAGATAATAACTAGTCCTCCAATGCTATCAGTTTCTATCAAGTTTTACTTGGCAACTGTTCTGGAGAATCCATTAGATCGATCTTGGAAACTCAAAGCTTTTGTGCCTTGTAATTAG

Protein sequence

MEFEIVAPGQNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSGETRPMSEGKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVPCN

Homology

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BLAST of CcUC04G069160 vs. NCBI nr
Match: WP_198843152.1 (hypothetical protein, partial [Aureibaculum sp. A20] >MBJ2176574.1 hypothetical protein [Aureibaculum sp. A20])

HSP 1 Score: 223.8 bits (569), Expect = 8.0e-55
Identity = 103/120 (85.83%), Postives = 114/120 (95.00%), Query Frame = 0


Query: 1   MEFEIVAPGQNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRV 60
           M+FE++APGQNDAL QNRC+PGGWEPI+NI DPYVQEMGRF+VMEHTHKLVGAS++FIRV
Sbjct: 5   MDFEVIAPGQNDALHQNRCKPGGWEPIKNIHDPYVQEMGRFSVMEHTHKLVGASMKFIRV 64

Query: 61  VSGETRPMSEGKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVPCN 120
           VSGETRPMSEGKEYRLVVEVAE IIT PPMLSVS+KFYLATVLE PLDRSW+L+AFV CN
Sbjct: 65  VSGETRPMSEGKEYRLVVEVAEHIITCPPMLSVSVKFYLATVLEKPLDRSWQLEAFVSCN 124

[Close]

BLAST of CcUC04G069160 vs. NCBI nr
Match: XP_038882290.1 (cysteine proteinase inhibitor 5-like [Benincasa hispida])

HSP 1 Score: 218.4 bits (555), Expect = 3.4e-53
Identity = 104/120 (86.67%), Postives = 113/120 (94.17%), Query Frame = 0


Query: 1   MEFEIVAPGQNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRV 60
           M++E+VAPGQNDALLQNRCEPG WEPI+NI DPYVQEMGRFAVMEHTHKLVGASL+FI V
Sbjct: 1   MDYEVVAPGQNDALLQNRCEPGCWEPIKNIHDPYVQEMGRFAVMEHTHKLVGASLKFICV 60

Query: 61  VSGETRPMSEGKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVPCN 120
           VSGETRPMS+GKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLE P+DRSW+L +FV  N
Sbjct: 61  VSGETRPMSQGKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLEKPIDRSWQLISFVSSN 120

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BLAST of CcUC04G069160 vs. NCBI nr
Match: WP_156424223.1 (hypothetical protein, partial [Myroides odoratus])

HSP 1 Score: 205.3 bits (521), Expect = 3.0e-49
Identity = 95/109 (87.16%), Postives = 104/109 (95.41%), Query Frame = 0


Query: 4   EIVAPGQNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSG 63
           E++APGQNDAL QNRC+PGGWEPI+NI DPYVQEMGRF+VMEHTHKLVGAS++FIRVVSG
Sbjct: 1   EVIAPGQNDALHQNRCKPGGWEPIKNIHDPYVQEMGRFSVMEHTHKLVGASMKFIRVVSG 60

Query: 64  ETRPMSEGKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLENPLDRSWK 113
           ETRPMSEGKEYRLVVEVAE IIT PPMLSVS+KFYLATVLE PLDRSW+
Sbjct: 61  ETRPMSEGKEYRLVVEVAEHIITCPPMLSVSVKFYLATVLEKPLDRSWQ 109

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BLAST of CcUC04G069160 vs. NCBI nr
Match: XP_022132557.1 (cysteine proteinase inhibitor 6-like [Momordica charantia])

HSP 1 Score: 131.7 bits (330), Expect = 4.1e-27
Identity = 71/116 (61.21%), Postives = 86/116 (74.14%), Query Frame = 0


Query: 6   VAPG--QNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSG 65
           VAP   QNDALL N  +PGGWEPI+NI DPYVQE+GRFAVMEHTH L GAS  FIRVVSG
Sbjct: 6   VAPAVQQNDALL-NVQKPGGWEPIKNINDPYVQELGRFAVMEHTHNLCGASYEFIRVVSG 65

Query: 66  ETRPMSEGKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVPC 120
           E++ +  GK+Y LV+EV E II   P +  SIKF+ A +L+   ++SW L +FVPC
Sbjct: 66  ESQVVG-GKKYCLVIEVKETIII--PSMPSSIKFFKAIMLDKAWEKSWALLSFVPC 117

[Close]

BLAST of CcUC04G069160 vs. NCBI nr
Match: XP_022132702.1 (cysteine proteinase inhibitor 5-like [Momordica charantia])

HSP 1 Score: 123.2 bits (308), Expect = 1.5e-24
Identity = 70/122 (57.38%), Postives = 87/122 (71.31%), Query Frame = 0


Query: 1   MEFEIVAPGQNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHK--LVGASLRFI 60
           M+   VAP QNDALL  +  PGGWEPI+NI+DPYVQEMGRFAVME   K   + AS+ FI
Sbjct: 1   MDSVAVAPAQNDALLHLKM-PGGWEPIKNIKDPYVQEMGRFAVMEQHPKPSELIASMEFI 60

Query: 61  RVVSGETRPMSEGKEYRLVVE--VAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAF 119
            VVSGE + ++ GKEYRLVVE  V E+II     +S SI+ Y+A VL+ P D+SWK+ +F
Sbjct: 61  GVVSGEIQLVAHGKEYRLVVEVRVTEKIIVHGCPVS-SIRLYVAVVLDKPSDKSWKVLSF 120

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BLAST of CcUC04G069160 vs. ExPASy Swiss-Prot
Match: Q41916 (Cysteine proteinase inhibitor 5 OS=Arabidopsis thaliana OX=3702 GN=CYS5 PE=2 SV=2)

HSP 1 Score: 61.2 bits (147), Expect = 9.0e-09
Identity = 41/99 (41.41%), Postives = 53/99 (53.54%), Query Frame = 0


Query: 22  GGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSGETRPMSEGKEYRLVVEVA 81
           GGW PI N+ DP V E+G FAV E+ +K   + L+F  VVSGET+ +S G  YRL V   
Sbjct: 29  GGWSPISNVTDPQVVEIGEFAVSEY-NKRSESGLKFETVVSGETQVVS-GTNYRLKVAAN 88

Query: 82  EQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVPCN 121
           +    S        K YLA V + P  +   L +F P N
Sbjct: 89  DGDGVS--------KNYLAIVWDKPWMKFRNLTSFEPAN 117

[Close]

BLAST of CcUC04G069160 vs. ExPASy Swiss-Prot
Match: Q10J94 (Cysteine proteinase inhibitor 8 OS=Oryza sativa subsp. japonica OX=39947 GN=Os03g0429000 PE=2 SV=1)

HSP 1 Score: 59.3 bits (142), Expect = 3.4e-08
Identity = 38/97 (39.18%), Postives = 53/97 (54.64%), Query Frame = 0


Query: 22  GGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSGETRPMSEGKEYRLVVEVA 81
           GGW PI ++ DP++QE+G +AV  H   L    LRF RV SGE + +S G  YRLVV  +
Sbjct: 33  GGWSPITDVGDPHIQELGGWAVERHA-SLSSDGLRFRRVTSGEQQVVS-GMNYRLVVSAS 92

Query: 82  EQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVP 119
           +     P   + S   Y+A V E     + +L +F P
Sbjct: 93  D-----PAGATAS---YVAVVYEQSWTNTRQLTSFKP 119

[Close]

BLAST of CcUC04G069160 vs. ExPASy Swiss-Prot
Match: Q10Q46 (Cysteine proteinase inhibitor 6 OS=Oryza sativa subsp. japonica OX=39947 GN=Os03g0210200 PE=3 SV=1)

HSP 1 Score: 57.4 bits (137), Expect = 1.3e-07
Identity = 28/61 (45.90%), Postives = 38/61 (62.30%), Query Frame = 0


Query: 21 PGGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSGETRPMSEGKEYRLVVEV 80
          P GW PI+NI DP++QE+GR+A+ E+        L F RV  GE + +S G  YRL +E 
Sbjct: 20 PRGWSPIKNIDDPHIQELGRWAITENNRVSPSDELTFHRVTGGEQQVVS-GMNYRLEIEA 79

Query: 81 A 82
          A
Sbjct: 80 A 79

[Close]

BLAST of CcUC04G069160 vs. ExPASy Swiss-Prot
Match: Q10Q47 (Putative cysteine proteinase inhibitor 7 OS=Oryza sativa subsp. japonica OX=39947 GN=Os03g0210100 PE=3 SV=1)

HSP 1 Score: 51.6 bits (122), Expect = 7.2e-06
Identity = 26/60 (43.33%), Postives = 37/60 (61.67%), Query Frame = 0


Query: 22 GGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSGETRPMSEGKEYRLVVEVA 81
          G WEP++ I DP+VQE+GR+AV E    +    L F RV  GE + ++ G  YRL +E +
Sbjct: 28 GSWEPVD-INDPHVQELGRWAVAEEDRGVAAGGLTFERVTDGEKQVVA-GVNYRLTLEAS 85

[Close]

BLAST of CcUC04G069160 vs. ExPASy TrEMBL
Match: A0A6J1BU61 (cysteine proteinase inhibitor 6-like OS=Momordica charantia OX=3673 GN=LOC111005388 PE=4 SV=1)

HSP 1 Score: 131.7 bits (330), Expect = 2.0e-27
Identity = 71/116 (61.21%), Postives = 86/116 (74.14%), Query Frame = 0


Query: 6   VAPG--QNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSG 65
           VAP   QNDALL N  +PGGWEPI+NI DPYVQE+GRFAVMEHTH L GAS  FIRVVSG
Sbjct: 6   VAPAVQQNDALL-NVQKPGGWEPIKNINDPYVQELGRFAVMEHTHNLCGASYEFIRVVSG 65

Query: 66  ETRPMSEGKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVPC 120
           E++ +  GK+Y LV+EV E II   P +  SIKF+ A +L+   ++SW L +FVPC
Sbjct: 66  ESQVVG-GKKYCLVIEVKETIII--PSMPSSIKFFKAIMLDKAWEKSWALLSFVPC 117

[Close]

BLAST of CcUC04G069160 vs. ExPASy TrEMBL
Match: A0A6J1BX23 (cysteine proteinase inhibitor 5-like OS=Momordica charantia OX=3673 GN=LOC111005501 PE=4 SV=1)

HSP 1 Score: 123.2 bits (308), Expect = 7.1e-25
Identity = 71/122 (58.20%), Postives = 89/122 (72.95%), Query Frame = 0


Query: 1   MEFEIVAPGQNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHKLVG--ASLRFI 60
           M+   VAP QNDA LQ+  +PGGWEPI+NI+DPYVQEMGRFAVME   K VG  AS+ F+
Sbjct: 32  MDSVAVAPAQNDA-LQHVKKPGGWEPIKNIKDPYVQEMGRFAVMEQP-KPVGLVASMEFV 91

Query: 61  RVVSGETRPMSEGKEYRLVVE--VAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAF 119
            VVSGE + ++ GKEYRLVVE  V E+II     +S SI+ Y+A VL+ P D+SWK+ +F
Sbjct: 92  GVVSGEIQLVAAGKEYRLVVEVKVTEKIIVHGCPVS-SIRLYVAVVLDKPSDKSWKVLSF 150

[Close]

BLAST of CcUC04G069160 vs. ExPASy TrEMBL
Match: A0A6J1BUK6 (cysteine proteinase inhibitor 5-like OS=Momordica charantia OX=3673 GN=LOC111005500 PE=4 SV=1)

HSP 1 Score: 123.2 bits (308), Expect = 7.1e-25
Identity = 70/122 (57.38%), Postives = 87/122 (71.31%), Query Frame = 0


Query: 1   MEFEIVAPGQNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHK--LVGASLRFI 60
           M+   VAP QNDALL  +  PGGWEPI+NI+DPYVQEMGRFAVME   K   + AS+ FI
Sbjct: 1   MDSVAVAPAQNDALLHLKM-PGGWEPIKNIKDPYVQEMGRFAVMEQHPKPSELIASMEFI 60

Query: 61  RVVSGETRPMSEGKEYRLVVE--VAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAF 119
            VVSGE + ++ GKEYRLVVE  V E+II     +S SI+ Y+A VL+ P D+SWK+ +F
Sbjct: 61  GVVSGEIQLVAHGKEYRLVVEVRVTEKIIVHGCPVS-SIRLYVAVVLDKPSDKSWKVLSF 120

[Close]

BLAST of CcUC04G069160 vs. ExPASy TrEMBL
Match: A0A6J1GCT2 (uncharacterized protein LOC111453027 OS=Cucurbita moschata OX=3662 GN=LOC111453027 PE=4 SV=1)

HSP 1 Score: 112.5 bits (280), Expect = 1.3e-21
Identity = 60/118 (50.85%), Postives = 80/118 (67.80%), Query Frame = 0


Query: 1   MEFEIVAPGQNDALLQNRCEPGGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRV 60
           M+ ++VA GQNDALL ++ +   WEP+ N+ D YVQE+GRFAVMEH H L G  + FI V
Sbjct: 1   MDLKVVAHGQNDALL-HKPKSCDWEPV-NVHDQYVQELGRFAVMEHVHGLCGVGMTFICV 60

Query: 61  VSGETRPMSEGKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVP 119
           VSGE++ + EGK+Y LVVEV E  +   P    +IK Y+A VL+    +SW+L AF P
Sbjct: 61  VSGESKMVCEGKKYSLVVEVKE--VPVLPHACPAIKLYVADVLDKQCAKSWELLAFKP 114

[Close]

BLAST of CcUC04G069160 vs. ExPASy TrEMBL
Match: A0A6J1FCN5 (cysteine proteinase inhibitor 8-like OS=Cucurbita moschata OX=3662 GN=LOC111442890 PE=4 SV=1)

HSP 1 Score: 104.0 bits (258), Expect = 4.5e-19
Identity = 53/114 (46.49%), Postives = 76/114 (66.67%), Query Frame = 0


Query: 7   APGQNDALLQNR--CEPGGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSGE 66
           APGQN+A+ Q++   E G W+PIE+I+DPYVQEMGRFAVMEH +  +G +L+FI V SGE
Sbjct: 8   APGQNEAMFQDKKGIEVGQWKPIEDIKDPYVQEMGRFAVMEHDNMRIGENLKFICVQSGE 67

Query: 67  TRPMSEGKEYRLVVEVAEQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVP 119
            + + EG  YRL+++ +  +  S          Y+A V + P +++WKL  FVP
Sbjct: 68  -KQVVEGMNYRLLIKASNHLGIS--------SLYMAIVYDKPWEKTWKLIEFVP 112

[Close]

BLAST of CcUC04G069160 vs. TAIR 10
Match: AT5G47550.1 (Cystatin/monellin superfamily protein )

HSP 1 Score: 61.2 bits (147), Expect = 6.4e-10
Identity = 41/99 (41.41%), Postives = 53/99 (53.54%), Query Frame = 0


Query: 22  GGWEPIENIRDPYVQEMGRFAVMEHTHKLVGASLRFIRVVSGETRPMSEGKEYRLVVEVA 81
           GGW PI N+ DP V E+G FAV E+ +K   + L+F  VVSGET+ +S G  YRL V   
Sbjct: 29  GGWSPISNVTDPQVVEIGEFAVSEY-NKRSESGLKFETVVSGETQVVS-GTNYRLKVAAN 88

Query: 82  EQIITSPPMLSVSIKFYLATVLENPLDRSWKLKAFVPCN 121
           +    S        K YLA V + P  +   L +F P N
Sbjct: 89  DGDGVS--------KNYLAIVWDKPWMKFRNLTSFEPAN 117

The following BLAST results are available for this feature:

BLAST of CcUC04G069160 vs. NCBI nr


Match Name	E-value	Identity	Description
WP_198843152.1	8.0e-55	85.83	hypothetical protein, partial [Aureibaculum sp. A20] >MBJ2176574.1 hypothetical ...	[more]
XP_038882290.1	3.4e-53	86.67	cysteine proteinase inhibitor 5-like [Benincasa hispida]	[more]
WP_156424223.1	3.0e-49	87.16	hypothetical protein, partial [Myroides odoratus]	[more]
XP_022132557.1	4.1e-27	61.21	cysteine proteinase inhibitor 6-like [Momordica charantia]	[more]
XP_022132702.1	1.5e-24	57.38	cysteine proteinase inhibitor 5-like [Momordica charantia]	[more]

BLAST of CcUC04G069160 vs. ExPASy Swiss-Prot


Match Name	E-value	Identity	Description
Q41916	9.0e-09	41.41	Cysteine proteinase inhibitor 5 OS=Arabidopsis thaliana OX=3702 GN=CYS5 PE=2 SV=...	[more]
Q10J94	3.4e-08	39.18	Cysteine proteinase inhibitor 8 OS=Oryza sativa subsp. japonica OX=39947 GN=Os03...	[more]
Q10Q46	1.3e-07	45.90	Cysteine proteinase inhibitor 6 OS=Oryza sativa subsp. japonica OX=39947 GN=Os03...	[more]
Q10Q47	7.2e-06	43.33	Putative cysteine proteinase inhibitor 7 OS=Oryza sativa subsp. japonica OX=3994...	[more]

BLAST of CcUC04G069160 vs. ExPASy TrEMBL


Match Name	E-value	Identity	Description
A0A6J1BU61	2.0e-27	61.21	cysteine proteinase inhibitor 6-like OS=Momordica charantia OX=3673 GN=LOC111005...	[more]
A0A6J1BX23	7.1e-25	58.20	cysteine proteinase inhibitor 5-like OS=Momordica charantia OX=3673 GN=LOC111005...	[more]
A0A6J1BUK6	7.1e-25	57.38	cysteine proteinase inhibitor 5-like OS=Momordica charantia OX=3673 GN=LOC111005...	[more]
A0A6J1GCT2	1.3e-21	50.85	uncharacterized protein LOC111453027 OS=Cucurbita moschata OX=3662 GN=LOC1114530...	[more]
A0A6J1FCN5	4.5e-19	46.49	cysteine proteinase inhibitor 8-like OS=Cucurbita moschata OX=3662 GN=LOC1114428...	[more]

BLAST of CcUC04G069160 vs. TAIR 10


Match Name	E-value	Identity	Description
AT5G47550.1	6.4e-10	41.41	Cystatin/monellin superfamily protein	[more]

InterPro

Analysis Name: InterPro Annotations of Watermelon (PI 537277) v1
Date Performed: 2022-01-31


IPR Term	IPR Description	Source	Source Term	Source Description	Alignment
None	No IPR available	GENE3D	3.10.450.10		coord: 18..119 e-value: 1.3E-17 score: 65.8
None	No IPR available	SUPERFAMILY	54403	Cystatin/monellin	coord: 22..118
IPR000010	Cystatin domain	PFAM	PF16845	SQAPI	coord: 27..117 e-value: 2.0E-11 score: 44.1
IPR000010	Cystatin domain	CDD	cd00042	CY	coord: 22..116 e-value: 7.61434E-7 score: 42.4612
IPR027214	Cystatin	PANTHER	PTHR47116	PHLOEM FILAMENT PROTEIN	coord: 19..118

Relationships

The following mRNA feature(s) are a part of this gene:


Feature Name	Unique Name	Type
CcUC04G069160.1	CcUC04G069160.1	mRNA

GO Annotation

GO Assignments

This gene is annotated with the following GO terms.

Category	Term Accession	Term Name
biological_process	GO:0010466	negative regulation of peptidase activity
molecular_function	GO:0004869	cysteine-type endopeptidase inhibitor activity